All your nitrates produced in the aquarium are coming from the marine life forms that are utilizing nitrogen foods. This of course includes the eel but don't forget the other life forms (of which there are many, which is another point).
The tank is a mix of invertebrates and fishes. To be honest, the tank should either be a FOWLR tank OR a reef tank. Too many fish in there to properly try and control the nitrates so that the corals are happy. An easy and properly functioning reef tank should have very few (if any) marine fishes.
You would have to work hard (many large water changes and frequently) and/or pay out a lot of money (nitrate and ammonia removers, reactors, deep sand bed, refugium, etc.) to try and control such a bioload of nitrate producers. Your setup seems to me to already be (normally or standard wise) maxed out to handle the 'normal' nitrate load (e.g., LR quantity is very good). [Although I don't know if that is the optimum skimmer or not.]



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